The hedgehog-rabbits in Eleum Loyce seem to be a reference to a philosophical concept called "The Hedgehog's Dilemma" (or "Porcupine's Paradox").
Hedgehogs are drawn together by a desire to nestle for warmth but are drawn apart by the pain their quills inflict upon each other.
Humans are also drawn together by a desire to feel the warmth of love, yet drawn apart by the pain they inevitably cause one another.
This simultaneous push-and-pull that Humanity engenders is much like the "Unity of Opposites" or "Logos" referenced by *Disparity* in the opening cinematic. It is especially noteworthy in Eleum Loyce because Alsanna's relationship was the only one typified by greater attraction than repulsion, rather than vice-versa.
I also believe that Dark is compared to "Logos" ("Word") by the Pus of Man, who spew ink and drop charcoal.
(Controversially, I also believe these enemies are a reference to the epithet, "snowbunnies," as a further reference to the books, "The White Goddess" and "V.," that I believe the trilogy draws inspiration from.
Another example is "Rose of Ariandel," being a reference to the folk song "Rose of Allendale" and "Yorshka" being a reference to the White Rose of Yorkshire.)
Sorry if this was a bit scattered. I just wanted to share a few ideas.